El complejo de especies Athyreus tribuliformis Felsche y descripciones de tres nuevas especies (Scarabaeoidea: Geotrupidae: Athyreini)

Resumo

The Athyreus tribuliformis Felsche complex is reviewed, all species are pictured, and a key to males is included. Three new species are described: Athyreus mouraensis Howden from Brazil, and A. bicornus Howden and A. larseni Howden from Peru.