Fuentes Medina, Paula Valeria, y Edgar Camero Rubio. 2009. «Coprophagous Beetles Fauna (Coleoptera: Scarabaeidae) in a Colombian Tropical Rain Forest». ENTOMOTROPICA 21 (3):133-43. https://saber.ucv.ve/ojs/index.php/rev_ento/article/view/7489.